Character design is based on the cubi of DMFA by missmab, who are characterized not by their wings but by having a second set of wings on their head. A third set of wings at the hips means that this particular cubi is a Clan Leader!

She's a rat/raccoon hybrid. Left her tail out of this particular image because I couldn't make it look good. (edit: almost a full year later, added her tail)

Ambrosia is the leader of the clan associated with the emotion of jealousy. She was born into clan Taun, but converted to Zezzuva's clan for unspecified reasons. When Zezzuva announced her "retirement" Ambrosia set out to aggressively breeding, bearing twenty-four children* over the course of twenty years. Seventy years later she and five of her sons announced their separation from the clan, and from that point on her sons were encouraged to breed as rapidly as possible and convert as many of the non-cubi mothers to cubi, by means of taking the children to SAIA immediately on their birth and telling the mothers that becoming a cubi is the only way they would ever see their children again.

After seven hundred years of this, and through intense brainwashing and various abuses both physical and psychological, Ambrosia was able to convince most of her female descendants and all but one of the converted mothers to mass-suicide in her name, and used the power their combined souls to ascend to tri-wing status. Her first order was for the male members of her clan to begin breeding more aggressively than ever.

Ambrosia insists that the emotion of jealousy is inherently different from the emotion of envy, but never explains HOW. As a clan leader she expects her words to be taken at face value, and doesn't like being questioned. Ever.

Clan quirk: Cubi born into Ambrosia's clan naturally have matching hair and eye colors.

*Cubi born into clan Zezzuva are always born twins, meaning Ambrosia was pregnant twelve times, with the births twenty months apart on average.

EDIT::: Storyline change: mothers and babies are both kidnapped, with the mothers being reduced to breeding stock if they want to live, and Ambrosia eating their souls once they've outlived their use to her. Shortly before the mass-suicide of her female descendants, Ambrosia murdered and soul-ate the remaining living non-cubi mothers.