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It comes from the attack on Toby's home planet. He's one of the few survivors of his race -- pulled from the rubble of a blast that killed many of the people he cared about. He's a refugee. In the previous comic, the word "stray" is used as a derogatory term for feline refugees.
Felix is happy out of optimistic and possibly naive world view
Toby is happy despite what he's suffered
Toby lashing out gets explained in the coming comics. At this time though, it can probably be assumed that someone laying a hand on his buddy had something to do with it
At the end of the day, Toby exemplifies one important key to happiness. Be greatful for what you have and count your blessings where you find them. Friends, food, shelter - not all are as fortunate as we are even for what little we may have.
And I love the coon. Kinda have to as I'm a coon too.