While not the ultimate antagonist of Yondard, he is effectively the main villain for most of Yondard.

One of the oldest Yondards, but also one of the strongest, and is fairly active. Among other things, he is the reason Fennia is missing a wing, and the initial reason the attack on Black Maw happened.

Don't let the fact that he's part rabbit fool you, most either fall in line or fall to his feet, and is involved with the backstory of all the main protagonists in some way.