Godzilla (ゴジラ Gojira) is a Titan who first appeared in the 2014 Legendary Pictures film Godzilla.

The Monsterverse Godzilla is the second incarnation of Godzilla to be featured in an American-made film, after the TriStar Godzilla from the 1998 GODZILLA film. A colossal apex predator who feeds on radiation, Godzilla has periodically surfaced throughout human history, worshiped as a god by a super-ancient civilization and inspiring some of humanity's mythology. In prehistoric times, Godzilla was involved in several ancient wars. He joined forces with Mothra to battle his alien nemesis, King Ghidorah, and succeeded in sealing the three-headed dragon in Antarctica. Godzilla also faced the Kong species during an ancient feud, locking away the tyrannical Skar King and his army of Great Apes in the depths of the Hollow Earth.

The multinational scientific agency Monarch became aware of Godzilla's existence in 1954; shortly thereafter, the United States attempted to kill him with the Castle Bravo hydrogen bomb test in Bikini Atoll. He became known to the public in 2014, when parasitic creatures from his time period dubbed MUTOs awakened and prepared to reproduce. He battled the male in Honolulu, then destroyed him and his female mate in a protracted duel in San Francisco. Later that year, he killed the MUTOs' parent, MUTO Prime, in Montana following a series of battles around the world. In 2015, he awakened in the Algerian Desert and traveled to the underground realm called Axis Mundi, where he soundly defeated the Ion Dragon.

When a group of eco-terrorists freed King Ghidorah from his icy tomb in 2019, Godzilla confronted the newly-awakened three-headed Titan, nearly defeating him in their second clash in the waters off Mexico before being interrupted by the U.S. military's Oxygen Destroyer. Godzilla retreated to his underwater lair in the Hollow Earth to heal, but his rejuvenation was accelerated by the sacrifice of Dr. Ishiro Serizawa, who set off a thermonuclear warhead in his vicinity. Godzilla, aided by Mothra, triumphed over Ghidorah in a final confrontation in Boston. The numerous Titans whom Ghidorah awakened submitted to Godzilla, accepting him as the King of the Monsters. Godzilla journeyed around the world to maintain balance among the these Titans and humanity for some time, but after locating a new home for himself, returned the Titans to their ancient rest.

In 2024, Godzilla sensed Apex Cybernetics' construction of Mechagodzilla, an artificial Titan meant to overthrow him, and began attacking the company's facilities, with humanity assuming he had turned against them. He intercepted a U.S. naval fleet transporting King Kong in the Tasman Sea, nearly killing the ape Titan. He swam to Hong Kong after he sensed Mechagodzilla being activated and drilled a hole into the Hollow Earth with his atomic breath, setting off a second battle with Kong where he again emerged triumphant. However, when he was overwhelmed by Mechagodzilla, Kong came to his aid and the two Titans reluctantly worked together to defeat their common enemy. After destroying Mechagodzilla, Godzilla and Kong went their separate ways, with Godzilla returning to the depths of the ocean.

In 2027, Godzilla hunted down and killed the arthropod Titan Scylla in Rome before she could consume enough nuclear energy to threaten him. While resting in the Colosseum, Godzilla sensed an Iwi signal warning of the return of the Skar King. Godzilla began gathering radiation to evolve into a powerful new form, attacking a nuclear power plant in France and killing Tiamat to claim her Norwegian lair's solar energy. Upon completing his evolution in the Arctic, Godzilla battled Kong again in Egypt after mistaking the ape's call for help as a challenge, nearly killing him before a newly reborn Mothra convinced him to join forces with Kong. Godzilla, Kong, Mothra, and Suko battled the Skar King and his enslaved ice Titan Shimo in the Hollow Earth and then in Rio de Janeiro, where they freed Shimo from the Skar King's control and killed the ape tyrant. After the victory, Godzilla returned to the Colosseum to rest.